Mourners hold pictures of Colonel Sayad Khodai, a member of Iran’s Islamic Revolution Guards Corps, in Tehran, Iran, May 24, 2022. Majid Asgaripour/WANA (West Asia News Agency) via REUTERS

Iran held a funeral procession on Tuesday in the centre of the capital Tehran for Revolutionary Guards Colonel Hassan Sayad Khodai, who was shot dead by two people on a motorcycle, and his commander vowed to avenge the attack.

State television showed crowds surrounding a truck carrying Khodai’s casket, wrapped in Iran’s flag and strewn with flowers. Mourners held portraits of Khodai, who was gunned down in broad daylight in front of his home in central Tehran on Sunday.

“Iran’s response to any threat or action will be harsh. But we will determine when and how it will be and in what circumstances. We will definitely take revenge on our enemies,” Revolutionary Guards commander Hossein Salami told reporters.

Iran has blamed such attacks on Israel. Foreign Ministry spokesperson Saeed Khatibzadeh said US support was making Israel more brazen.

“There is no doubt that the overt and covert support of …the United States plays an important role in increasing the audacity of the occupying regime (Israel),” Foreign Ministry spokesperson Saeed Khatibzadeh told state media.

Separately, state television said the Guards had arrested members of a network of “thugs” recruited by Israeli intelligence to carry out sabotage and attacks in Iran.

The Israeli Prime Minister’s office, which oversees intelligence agency Mossad, has declined to comment on the events in Tehran.

Israeli media said Khodai headed a unit of the Quds Force — the Revolutionary Guards’ overseas arm — planning attacks on Israelis abroad.

Khodai was a “defender of the shrines,” Iranian state media said on Sunday, referring to military personnel or advisers who Iran says fight on its behalf to protect Shi’ite sites in Iraq or Syria against groups such as Islamic State.

The forces have played a key role in backing Syrian President Bashar al-Assad, Tehran’s ally.